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Wasps like to kill the queen if they can get in the hive.The damage looks like hornets.I've battled with them 2 years in a row.Finally found their hives and destroyed them.If you see grey stuff on the edges inside the void I would say wasp.In my area wasp seem to be a problem in the spring and the hornets are a problem in the spring.Probably dependent on which has a higher population or who's preferred food source is available.Hornets work in a helical pattern kind of looks like what is going on on your comb.Yes hornets like to pinch all castes in half and cause mass destruction.Wasp seem to target the queen maybe pinch just a few workers to get to her.That's what I've witnessed firsthand.Wasp got one of my queens this spring,didn't pinch any workers but put the hit on the queen.:pinch:
 

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Most surely sounds like hornets.The grey would be noticeable.It was so cold this year I'm sure more so up your way than mine.It was an unusally cold winter in my area also.Bees with heads in the cell signifies starvation.I've never known bees to display cowardice and hide from an intruder.With several polar vortexs this winter it was difficult for your bees to make the jump for feed.That's more than likely what happened.Good luck with your studies:applause:Me after locating a hornets nest:banana:
 
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