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Interesting read. I noticed that this was done in 1996. It sort of coincides with the narrower frame theory that a warmer brood nest helps to reduce varroa. However, I don't agree with much that has been said. Varroa does quite well in the subtropics of Hawaii where they have been present for about 10 years. Each year seems to be different in how the varroa manifests itself. My opinion is that brood diseases related to varroa seem to make the queen less productive while the bees often fail to raise a new queen in response. Eventually the amount of pollen in the hive builds up (assuming good moisture conditions) and then the small hive beetles take over and destroy the hive. Dead hive leads to mite bomb and so it goes.
 
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