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Odemer (2020) Reproductive capacity of varroa destructor in four different honey bee subspecies https://www.sciencedirect.com/science/article/pii/S1319562X19301640?via=ihub
speaks to just how fast traits are lost with outcrosingFor the F2 generation of the surviving population from Gotland however, we had expected a different outcome. The Gotland bees have developed an apparent reduced mite reproductive success trait that is either inheritable from paternal, maternal or both sides in the F1 generation (Locke, 2016b). Our results provide evidence that this trait seems to fade out by further generational change, once more making the colonies susceptible to Varroosis.