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At least around here, Greater Waxworm larvae will tunnel through frames in much the same manner as the photos you posted.

They will sometimes be on the side or bottom bars, but are most often in the corners where top and sidebars meet and through the topbar as shown in your photos.

Again around here, when put back in a strong colony they will often propolize the holes shut, so the frames are almost as good as new. ;)
 
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