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I would say they came that way, looks like carpenter ants that were in the tree when harvested.
 

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At least around here, Greater Waxworm larvae will tunnel through frames in much the same manner as the photos you posted.

They will sometimes be on the side or bottom bars, but are most often in the corners where top and sidebars meet and through the topbar as shown in your photos.

Again around here, when put back in a strong colony they will often propolize the holes shut, so the frames are almost as good as new. ;)
 

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Mutts and Carniolans in Langs.
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North Georgia USA Zone 7b
At least around here, Greater Waxworm larvae will tunnel through frames in much the same manner as the photos you posted.
Do they also make the shallow oval 'pockets' under their cocoons? All I have seen and no actual through holes like in the pictures above. Distinction between greater and lessor waxworm?
 

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That is a classic sign of beetle damage. If you look you'll see the frass lining the tunnels, and there's a beetle pupa in the top tunnel. I wouldn't worry about it as I've seen the, as well.
 
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