The cluster has to move throughout the winter to stay connected to stores. Usually where the main portion of bees are when it's warm is where they will cluster when it gets cold. That location usually has a core of empty cells that the bees can climb into. That location changes through the winter as the consume the stores so the cluster stays where the edge of the cluster is in contact with stores. If they fail to do this (and they do often enough to be frustrating) then they starve in place. Usually this is caused by them not being willing to move off of brood.
>Thanks. If they had stores in two boxes, do you think the cluster could extend from one box to the other?
The cluster is usually roughly spherical. That sphere, as it moves up, is often bridging the gap between the boxes, but there is a tendency for the bees to try to move up to the cluster rather than stay in the gap when there are only a few bees below, so once it gets 2/3 of the way or so across the gap they tend to all migrate up to the next box, at least while it's cold. They may spread back out during a warm spell and span the gap again.
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