Less male mites are able to mate:
Reproduction of Varroa destructor in South African honey bees: does cell space influence Varroa male survivorship, Stephen J. MARTIN, Per KRYGER :
http://www.apidologie.org/index.php...29&url=/articles/apido/pdf/2002/01/Martin.pdf
Less mite offspring:
The influence of brood comb cell size on the reproductive behavior of the ectoparasitic mite Varroa destructor in Africanized honey bee colonies Giancarlo A. Piccirillo1 and D. De Jong :
http://funpecrp.com.br/gmr/year2003/vol1-2/gmr0057_full_text.htm
Smaller cell size significantly contributes to reduced reproductive success of Varroa: Cell size and Varroa destructor mite infestations in susceptible and naturally-surviving honeybee (Apis mellifera) colonies :
https://link.springer.com/article/1...9aWvRQWlImQcfRifvq1XzsHHytlZspO5f_z5B1l8IrKLg
Survival of a Commercial Beekeeper in Norway, Hans Otto Johnsen :
https://www.beesource.com/point-of-...survival-of-a-commercial-beekeeper-in-norway/
Trial of HoneySuperCell® Small Cell Combs, Randy Oliver :
http://scientificbeekeeping.com/trial-of-honeysupercell-small-cell-combs/
Effects of Comb Cell Diameter on Parasitic Mite Infestations in Honey Bee Colonies, Eric Erickson Jr. :
https://beesource.com/point-of-view...itic-mite-infestations-in-honey-bee-colonies/
The workers live longer:
On small cell bee longevity: Life span of worker honeybees reared in colonies kept on small-cell combs, Krzysztof Olszewski, Grzegorz Borsuk, Jerzy Paleolog, Aneta Strache :
http://medycynawet.edu.pl/images/stories/pdf/pdf2014/122014/201412777780.pdf
The pupation cycle is one to two days shorter by my measurement and that of Huber and Dzierzon:
On Shortened Pupation: A model of the mite parasite, Varroa destructor, on honeybees (Apis mellifera) to investigate parameters important to mite population growth, D Wilkinson, G.C Smith :
http://www.sciencedirect.com/science/article/pii/S0304380001004409