I love the discussion of so many heavyweights in the ring. This thread is a fun read, and it reminds of something my dad once said.
"I went to see a fight and a hockey match broke out!"
From a retired poker player's ("Know when to walk away, know when to run..") viewpoint, somewhat versed in statistics, genetics, lower division mathematics (3rd semester calculus completed, differential calculus started but not completed, although I tutored that, physics, chemistry, and many other subjects for 15+ years) my bets are that mite mauling is the single best trait to promote. Add a brood break - they usually survive. Add allogrooming - they usually survive. Delve deeply into the WHY of heavy honey production and analyzing one's own methods, and you are probably breeding bees that serve your purposes.
CHANGE YOUR METHODS TO 5.1 mm CELL SIZE COMBINED WITH NARROW (1.240 INCH WIDE) FRAMES to take better advantage of the main Spring nectar / pollen flow, and you likely one of the bigger beekeepers in the neighborhood.
Until any of us can prove we are right and that God does or does not exist, we may cite some bitchin' studies, but we are, like it or not, are a bunch of agnostics. All I have are opinions, backed up with 17% more turbo charged loadabunk than JWC, msl, Bigfoot, or Oldtimer, so there!