When you allow the bees to build comb in foundationless frames you'll find that they build mainly worker size comb in the brood nest and mainly drone size comb outside the brood nest where it's used for honey storage for the majority of the year. This is especially noticeable in a horizontal hive with an entrance at one end.
In a wild hive this boundary between the brood nest and honey storage comb is reached about the time the hive would start raising drones.
I find it interesting that beekeeping with it's use of foundation has not promoted using worker size foundation only in the brood nest and drone size comb only for honey supers.
Doing this would also act like a queen excluder as the quuen would only lay a certain amount of drone in the box directly above the brood nest. Above that would be only honey.
Also, pollen is less likely to be stored in drone size cells.