1. A cold-blooded animal is
one that has a body temperature below 80 degrees F., and that
takes on the temperature of the air, water, or other element
in which it lives. One bee, or a few bees, do take on the temperature
of the air around them and cannot protect themselves against
the loss of heat or cold.
2. A warm-blooded animal is one having a relatively high and
constant body temperature relatively independent of the surrounding
environment. The bee cluster can keep itself warm against a temperature
of 100 degrees F below zero or cool against a temperature of
over 135 degrees F by metabolic activity mimicking warm-bloodedness
by working together as a whole harmonious unit to provide an
optimum and constant body temperature relatively independent
of the harsh surrounding conditions of temperature and humidity.
3. With an internal ambient temperature of approximately 106
degrees F both bees and brood die without some measures of heat
regulation.
4. When the ambient temperature inside the hive drops to 45 degrees
F, bees normally cease work, cluster loosely, and maintain the
cluster temperature at 57 - 58 degrees F.
5. The cluster is most nearly dormant at 57 - 58 degrees F which
still allows the bees to be able to break cluster and move to
a new store of honey when all within the cluster has been consumed.
6. Honeybee clusters generate 12 - 13 degrees F heat by their
normal and natural bodily metabolism or activity incidental to
living.
7. The brood rearing temperature is approximately 93 degrees
F to stimulate both the queen to lay eggs and the worker bees
to feed and care for larvae.
8. Once the brood rearing has begun, bees must generate whatever
heat it takes to maintain the brood nest temperature at approximately
93 degrees F until the brood emerges.
9. If the temperature of the outside air rises to 90 degrees
F or higher, bees normally carry water into the hive, evaporating
it by forced air circulation and thus removing the excess heat
from the hive (Evaporation of water cools the hive because the
specific heat of water is more than 4 times that of air).
10. Pure hybridization occurs where hot-weather bees (yellow)
and cold-weather bees (black/brown) come together naturally by
either latitude or altitude with a mean monthly temperature of
75 degrees F. (Note: Now refer back to Saga #8 picturing in your
mind the world as a basic map of honeybee thermal/cell size zones
based on composites of hot and cold land area masses. Read and
recap here, then proceed on.)
11. As the inside ambient temperature approaches and/or exceeds
both 45 degrees F and 106 degrees F small black bees approach
the breeding condition of thelytoky (Have not been able to accomplish
with either yellow-mix or large dark castes.)
12. Humidity in the brood chamber should be about 60% relative
humidity, while in the supers where the honey is being ripened
it should be 10% relative humidity.
OTHER BASIC GUIDLINES FOR BEE BREEDING
(Note: Now refer back to Saga #8 again picturing in your mind
the world as a basic map of honeybee thermal/cell size zones
based on composites of hot and cold land area masses. Read and
recap here, then proceed on.):
1. Dark (brown/black) cold-weather bees exist naturally below
30 degrees latitude where higher altitudes permit.
2. (Yellow) hot-weather bees exist naturally above 30 degrees
latitude where warm thermal areas permit.
3. Small caste races/strains of hot-weather (yellow) bees exist
at the Equator and large caste races/strains of cold-weather
(brown/black) bees exist as they approach the poles.
4. As all races/strains of bees advance towards temperature transition-zones
at near 30 degrees latitude, hot-weather (yellow) bees hybridize
more (larger), while cold-weather (brown/black) bees hybridize
less (stay more towards pure races/strains and smaller).
5. Nature breeds constantly and constant when all optimum basic
evolutionary needs are met i.e. water, food, shelter, and temperature.
6. Mongrel hybridization (man made) is not an evolutionary progression
for it separates when artificial stimuli are removed i.e. inappropriate
artificial bigger comb size, surrogate geographic areas, and
forced climatic breeding.
7. Nature breeds evolutionary change that is progressive, retrogressive,
or cloned, when race/strain survivability is at stake.
8. Each race/strain of honeybees has its own separate breeding
cycle in Nature providing, an evolution separate from all others,
enabling it to exist.
9. Large caste bees on a natural system equate with: 1) fewer
bees per brood comb, 2) slower developmental time, and 3) slower
mating flight speed.
10. Small caste bees on a natural system equate with: 1) more
bees per brood comb, 2) faster developmental time, and 3) faster
mating flight speed.
11. Drones take mating flights only on days when bees are able
to break cluster and fly outside.
In queen rearing, not the outside air temperature itself is the
focal point which beekeepers must consider, but the temperature
of the skin surface of the artificially boxed hive where exposed
to the sun or the chill-factor of cold winds, which may reach
135 degrees F or 100 degrees F below zero, or even more depending
upon latitude and altitude, and time of the year. This heat or
cold passes through the wall/entrance of the hive to its interior,
thus increasing or decreasing it to far above or below the outside
temperature. Beekeepers seriously breeding bees can help colonies
thermoregulate by maintaining tight and painted equipment, and
leaving full frames of honey surrounding the brood nests to act
as insulation against extremes of cold and heat.
By natural metabolic cluster reactions, honeybees thermodynamically
overcome these effects of unfavorable weather conditions within
the hive during cold winters and hot summers. However, to bees,
the temperature of the skin surface of the artificial box is
a trigger mechanism to which they must react, to average the
maximum and minimum temperatures of each day. Day in and day
out, bees must manipulate natural weather conditions to approach
and provide optimum mean temperature conditions for brood rearing
and colony survival.
An ambient temperature lower than about 80 degrees F inside the
colony results in one of two things. Either the brood rearing
within the colony decreases and cuts back or, if seasonal conditions
cause the bees to react favorably (fresh pollen and/or nectar
coming in), the bees will increase their metabolic activity and
produce the necessary heat to offset any short-term decrease
in temperature, adding a minimum of 12 - 13 degrees F of their
own body heat to raise brood, if there is a supply of pollen
and reserve honey stored. As soon as the brood rearing temperature
of 93 degrees F is reached, the queen begins to lay eggs and
the brood is reared and cared for by the colony.
In spring, when most beekeepers think of rearing queens, they
think of progressive breeding techniques, waiting until colonies
produce sufficient drones and nurse bees before beginning their
queen rearing. MANY WRONGLY BELIEVE THAT HYBRIDIZATION IS PROGRESSIVE
BREEDING. IT IS NOT! In today's world, HYBRIDIZATION IS FOR THE
MOST PART MONGREL BREEDING THAT PRODUCES ONLY A SHORT BURST OF
HYBRID VIGOR AND THEN QUICKLY FALLS APART WITH EACH SUCCEEDING
GENERATION.
For most beekeepers, there should be no breeding from hybrids
since it is beyond most beekeepers to control it (We will talk
about this control and the field mechanics involved before we
end the bee breeding portion of this saga.). The final result
is nearly always total mongrelization of local area bee stocks
and an uncontrolled mixture of overly aggressive honeybees which
makes beekeeping more and more impossible in today's urbanizing
world. (Note: Could this be an underlying causative effect giving
rise to a myth of "Africanized Killer Bees" as being
uncontrollable, when the real cause is probably mongrelization
with beekeepers mixing uncontrolled variances of races/strains
of honeybees together, as each one uses the type of honeybees
he/she prefers?)
In a long-term stock improvement program, artificial insemination
and various closed-population breeding methods should be avoided,
as they lead to severe inbreeding, resulting in poor brood patterns,
poor product averages, weak winter cluster carry-over, and colony
collapse over a period of 20 - 30 years. This is not to say however,
that artificial insemination methods for honeybee queens does
not have its place in bee breeding, but the technique is definitely
over used in today's world. In skilled hands, the technique of
artificial insemination can save many years work in development
of properly field-managed stock lines of several hundred colonies,
when used in conjunction with a modified open-mating system.
Nature breeds evolutionary changes that are progressive, retrogressive,
or cloned, when race/strain survivability is at stake. To accomplish
either of the three, beekeepers must remember that all breeding
begins with the selection of notable breeding stock of above
average overall colony performance. Beekeepers should look for
and select honeybee breeder colonies based on a whole-bee theory
of field characteristics. To do anything else will, in the long-term,
doom the breeding program to problems and necessitate retrogression
before being able to proceed further. (Note: We will talk about
a whole-bee theory of field characteristic selection and the
field mechanics involved before we end the bee breeding portion
of this saga.)
Retrogression in a bee hive is not a simple process. We have
talked about cell size retrogression and what it involves in
physically sizing honeybees back down to natural feral sizing
for control of all acarapis mites and their accompanying secondary
diseases. This necessary process sets the stage for bee breeding
as survivability and variability are achieved. But, just what
is progressive breeding, retrogressive breeding (not to be confused
with retrogression relative to size), and cloning (thelytoky)
as pertains to breeding honeybees?
PROGRESSIVE BREEDING: Is the production of uniform progeny within
the framework of a fully naturalized breeding program which will
true breed and the results of which can only be obtained from
uniformly bred colonies. Permanent results can only be achieved
by the use of naturally occurring races/strains of honeybees.
Since a bee by any other name is still a bee, then beekeepers
must use individual or combinations of large or small caste races/strains
of hot (yellow) or cold-weather (brown/black) bees to accomplish
this.
Artificial hybrids may then be created by mimicking natural hybridization,
when two of these races/strains are assimilated. Nature does
not produce complex mongrels. Nature transitions in and out from
one race/strain to another, with a brief transition-zone between,
that is a mixture of each, while always maintaining compatibility
to localized geography and climatic thermodynamics.
RETROGRESSIVE BREEDING: Is the reversal of either natural or
artificial hybridized combinations of large or small caste races/strains
of hot or cold-weather bees, resulting in the production of uniform
progeny within the framework of a fully naturalized breeding
program, which will then result in each separation achieved,
breeding true to their own hot or cold-weather characteristics
and large or small caste delineations.
Results can only be achieved by the use of stress-breeding at
either the beginning or the end of the selected race/strain breeding
cycles where no overlap occurs, one projected breeder-cycle to
the other(s). Artificial races/strains can then be created by
mimicking natural races/strains where complex mongrelization
has taken place, to gain uniformity of characteristics then necessary
for the advancement of desirable traits i.e. gentleness and production.
CLONING (THELYTOKY): Is the holding constant of race/strain genetics
from one generation to the next naturally or by artificially
increasing the propensity of worker bees to lay viable brood,
to raise queens as an alternate survival system to supplement
normal queen mating in case the virgin queen is lost during the
mating flight.
Results can only be achieved by using severe stress-breeding,
by using the temperature outside, the beginning or the end of
selected race/strain breeding cycles where no overlap occurs,
one projected breeder cycle to the other(s).
It is a short-duration phenomenon initiated by extreme stress
to allow perpetuation of species, until the first available normal
mating can be accomplished, to allow, the colony to permanently
requeen itself in the normal manner of mating. (Note: Simply
put, thelytoky is a natural state in Nature found in low incidence
in Italian bees and in higher incidence in natural sized small
black/brown honeybees that allows for workerbees to be self-fertile
for short periods of time so when hives go queenless during hot
dearth summers or extremely cold winters, they can requeen themselves
with a temporary queen to carry on until season is such that
the colony can requeen themselves naturally again.)
--
Signed: Dee A. Lusby, Tucson, Arizona, USA, 1-520-748-0542
Email Address: deealusby1@aol.com |