I was talking with Walter Wright about seasonal bee behavior in Langstroth equipment. He has been watching and writing about it for almost a decade.
Walter has also removed feral hives from between floor joists. He noted that the broodnest structure was alot like the vertical Lang but just tipped on it's side.
I hadn't thought about it that way. But that's what I've seen in my tbh and pictures of vertically oriented feral hives.