I was measuring the comb spacing on natural comb. These were the bees that were mostly full sized that moved into a feeder. It was a small colony and it quickly regressed as indicated by the cell sizes. The first combs in the center were about 5.1mm The next combs beside that were about 5.0mm and the next were about 4.9mm and the last were about 4.8mm. I think it was small enough that some of the bees emerged from the first comb by the time they were building the next combs.
The spacing on the combs varied. The combs ran dianoly at almost exactly 45 degrees from the side wall, until the ends where they rotated until they were almost parallel to the side wall.
The spacings varied from 1 1/8" to 1 5/8" between the centers of the combs. The most common spacing was 1 1/4"