In my short experience, I've observed that the bees kinda do what they want, regardless of how we try to trick them. I have some cordovans that have really messed up a couple of frames with tunnel, ladder and bridge comb. They have smaller cells and large drone cells all mixed up together. I'm wondering if orderly cell building is a trait you can select for?
I have some darker bees that have started drawing cells nicely on new wax foundation. Seems like they do alright in the center of the comb, get sorta confused when they get near the edges.
Banjos and bees... how sweet it is!