Vorroa mite winter behaviours

[Ian]

Does anyone have any information on the winter behaviour and habbits of the vorroa mite in a broodless wintering hive? Links or comments would be appreciated.
Are they quite dosile, just hanging on, or do they move about frequently, or do they dig right into the bee hard to reach and hard to stir?

Thanks

[Ian]

Here is a small bit of some interesting information found,

http://journals.cambridge.org/action/displayAbstract?fromPage=online&aid=23647

Anyother links or comments welcome,